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Research and Development Arrangements (Tables)
 9 Months Ended
Sep. 30, 2018
Astellas [Member]  
Summary of Research and Development Revenue

Revenue from Astellas included (in thousands):

 

 

Three Months Ended

 

 

Nine Months Ended

 

 

 

September 30, 2018

 

 

September 30, 2017

 

 

September 30, 2018

 

 

September 30, 2017

 

Research and development revenues

 

$

8,526

 

 

$

2,112

 

 

$

16,791

 

 

$

8,810

 

License revenues

 

 

1,915

 

 

 

318

 

 

 

5,133

 

 

 

6,707

 

Total Revenue from Astellas

 

$

10,441

 

 

$

2,430

 

 

$

21,924

 

 

$

15,517