SEC FORM 4

SEC Form 4

FORM 4

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP

Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934
or Section 30(h) of the Investment Company Act of 1940

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Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).

1. Name and Address of Reporting Person^*

KERR BARBARA J

(Last)

(First)

(Middle)

APPLERA CORP - APPLIED BIOSYSTEMS GROUP

850 LINCOLN CENTRE DRIVE

(Street)

FOSTER CITY

94404

(City)

(State)

(Zip)

2. Issuer Name and Ticker or Trading Symbol
APPLERA CORP [ ABI/CRA ]

5. Relationship of Reporting Person(s) to Issuer
(Check all applicable)

	Director	10% Owner
X	Officer (give title below)	Other (specify below)
	Vice President-Human Resourses

3. Date of Earliest Transaction (Month/Day/Year)
11/01/2007

4. If Amendment, Date of Original Filed (Month/Day/Year)

6. Individual or Joint/Group Filing (Check Applicable Line)

X	Form filed by One Reporting Person
	Form filed by More than One Reporting Person

Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned
1. Title of Security (Instr. 3)	2. Transaction Date (Month/Day/Year)	2A. Deemed Execution Date, if any (Month/Day/Year)	3. Transaction Code (Instr. 8)		4. Securities Acquired (A) or Disposed Of (D) (Instr. 3, 4 and 5)			5. Amount of Securities Beneficially Owned Following Reported Transaction(s) (Instr. 3 and 4)	6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	7. Nature of Indirect Beneficial Ownership (Instr. 4)
1. Title of Security (Instr. 3)	2. Transaction Date (Month/Day/Year)	2A. Deemed Execution Date, if any (Month/Day/Year)	Code	V	Amount	(A) or (D)	Price		6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	7. Nature of Indirect Beneficial Ownership (Instr. 4)
Applied Biosystems Group Common Stock	11/01/2007		M		8,364	A	$15.54	23,925.7177⁽¹⁾	D
Applied Biosystems Group Common Stock	11/01/2007		M		9,936	A	$20.185	33,861.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		100	D	$37.34	33,761.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		100	D	$37.33	33,661.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		100	D	$37.32	33,561.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		200	D	$37.26	33,361.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		800	D	$37.2	32,561.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		200	D	$37.14	32,361.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		500	D	$37.13	31,861.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		100	D	$37.12	31,761.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		100	D	$37.11	31,661.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		100	D	$37.08	31,561.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		1,200	D	$37	30,361.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		600	D	$36.99	29,761.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		900	D	$36.98	28,861.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		600	D	$36.97	28,261.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		200	D	$36.96	28,061.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		500	D	$36.95	27,561.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		100	D	$36.94	27,461.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		100	D	$36.9	27,361.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		300	D	$36.89	27,061.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		200	D	$36.88	26,861.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		400	D	$36.87	26,461.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		500	D	$36.86	25,961.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		800	D	$36.85	25,161.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		800	D	$36.84	24,361.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		700	D	$36.83	23,661.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		800	D	$36.79	22,861.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		1,000	D	$36.78	21,861.7177	D
Applied Biosystems Group Common Stock	11/01/2007		S⁽²⁾		3,300	D	$36.77	18,561.7177	D

Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned (e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivative Security (Instr. 3)	2. Conversion or Exercise Price of Derivative Security	3. Transaction Date (Month/Day/Year)	3A. Deemed Execution Date, if any (Month/Day/Year)	4. Transaction Code (Instr. 8)		5. Number of Derivative Securities Acquired (A) or Disposed of (D) (Instr. 3, 4 and 5)		6. Date Exercisable and Expiration Date (Month/Day/Year)		7. Title and Amount of Securities Underlying Derivative Security (Instr. 3 and 4)		8. Price of Derivative Security (Instr. 5)	9. Number of derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4)	10. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	11. Nature of Indirect Beneficial Ownership (Instr. 4)
1. Title of Derivative Security (Instr. 3)	2. Conversion or Exercise Price of Derivative Security	3. Transaction Date (Month/Day/Year)	3A. Deemed Execution Date, if any (Month/Day/Year)	Code	V	(A)	(D)	Date Exercisable	Expiration Date	Title	Amount or Number of Shares	8. Price of Derivative Security (Instr. 5)		10. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	11. Nature of Indirect Beneficial Ownership (Instr. 4)
Applied Biosystems Group Employee Stock Options-Right to Buy	$15.54	11/01/2007		M			8,364	⁽³⁾	03/24/2013	Applied Biosystems Group Common Stock	8,364	$0	41,336	D
Applied Biosystems Group Employee Stock Options-Right to Buy	$20.185	11/01/2007		M			9,936	⁽⁴⁾	06/17/2014	Applied Biosystems Group Common Stock	9,936	$0	47,664	D

Explanation of Responses:

1. Includes 126.1394 shares of Applied Biosystems Group Common Stock purchased in September 2007 under the issuer's employee stock purchase plan.

2. Reflects sale by independent third-party administrator under the issuer's Insider Diversification Program.

3. These options originally became exercisable in four equal annual installments, commencing March 24, 2004. Effective January 20, 2005, the vesting of all of these options was accelerated. However, shares received upon the exercise of accelerated options were subject to a restriction on transfer (covering sales, gifts, pledges, and any other method of disposition) that expired on the original vesting dates of those options.

4. These options originally became exercisable in four equal annual installments, commencing June 17, 2005. Effective January 20, 2005, the vesting of all of these options was accelerated. However, shares received upon the exercise of accelerated options are subject to a restriction on transfer (covering sales, gifts, pledges, and any other method of disposition) that will remain in effect until the earlier of the original vesting dates of those options or termination of the reporting person's employment.

Remarks:

This is the first of two forms being filed by the reporting person on November 5, 2007.

	/s/ Thomas P. Livingston, Attorney-In-Fact for Barbara J. Kerr	11/05/2007
	** Signature of Reporting Person	Date
Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
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