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<SEC-DOCUMENT>0001209191-09-051984.txt : 20091110
<SEC-HEADER>0001209191-09-051984.hdr.sgml : 20091110
<ACCEPTANCE-DATETIME>20091110180014
ACCESSION NUMBER:		0001209191-09-051984
CONFORMED SUBMISSION TYPE:	4
PUBLIC DOCUMENT COUNT:		1
CONFORMED PERIOD OF REPORT:	20091110
FILED AS OF DATE:		20091110
DATE AS OF CHANGE:		20091110

ISSUER:		

	COMPANY DATA:	
		COMPANY CONFORMED NAME:			Ancestry.com Inc.
		CENTRAL INDEX KEY:			0001469433
		STANDARD INDUSTRIAL CLASSIFICATION:	SERVICES-COMPUTER PROCESSING & DATA PREPARATION [7374]
		IRS NUMBER:				261235962
		STATE OF INCORPORATION:			DE
		FISCAL YEAR END:			1231

	BUSINESS ADDRESS:	
		STREET 1:		360 WEST 4800 NORTH
		CITY:			PROVO
		STATE:			UT
		ZIP:			84604
		BUSINESS PHONE:		801-705-7000

	MAIL ADDRESS:	
		STREET 1:		360 WEST 4800 NORTH
		CITY:			PROVO
		STATE:			UT
		ZIP:			84604

REPORTING-OWNER:	

	OWNER DATA:	
		COMPANY CONFORMED NAME:			PARKER VICTOR E JR
		CENTRAL INDEX KEY:			0001166804

	FILING VALUES:
		FORM TYPE:		4
		SEC ACT:		1934 Act
		SEC FILE NUMBER:	001-34518
		FILM NUMBER:		091173162

	MAIL ADDRESS:	
		STREET 1:		333 MIDDLEFIELD ROAD,
		STREET 2:		SUITE 200
		CITY:			MENLO PARK
		STATE:			CA
		ZIP:			94025
</SEC-HEADER>
<DOCUMENT>
<TYPE>4
<SEQUENCE>1
<FILENAME>doc4.xml
<DESCRIPTION>FORM 4 SUBMISSION
<TEXT>
<XML>
<?xml version="1.0"?>
<ownershipDocument>

    <schemaVersion>X0303</schemaVersion>

    <documentType>4</documentType>

    <periodOfReport>2009-11-10</periodOfReport>

    <notSubjectToSection16>0</notSubjectToSection16>

    <issuer>
        <issuerCik>0001469433</issuerCik>
        <issuerName>Ancestry.com Inc.</issuerName>
        <issuerTradingSymbol>ACOM</issuerTradingSymbol>
    </issuer>

    <reportingOwner>
        <reportingOwnerId>
            <rptOwnerCik>0001166804</rptOwnerCik>
            <rptOwnerName>PARKER VICTOR E JR</rptOwnerName>
        </reportingOwnerId>
        <reportingOwnerAddress>
            <rptOwnerStreet1>360 WEST 4800 NORTH</rptOwnerStreet1>
            <rptOwnerStreet2></rptOwnerStreet2>
            <rptOwnerCity>PROVO</rptOwnerCity>
            <rptOwnerState>UT</rptOwnerState>
            <rptOwnerZipCode>84604</rptOwnerZipCode>
            <rptOwnerStateDescription></rptOwnerStateDescription>
        </reportingOwnerAddress>
        <reportingOwnerRelationship>
            <isDirector>1</isDirector>
            <isOfficer>0</isOfficer>
            <isTenPercentOwner>0</isTenPercentOwner>
            <isOther>0</isOther>
        </reportingOwnerRelationship>
    </reportingOwner>

    <nonDerivativeTable>
        <nonDerivativeTransaction>
            <securityTitle>
                <value>Common Stock, $0.001 par value</value>
            </securityTitle>
            <transactionDate>
                <value>2009-11-10</value>
            </transactionDate>
            <deemedExecutionDate></deemedExecutionDate>
            <transactionCoding>
                <transactionFormType>4</transactionFormType>
                <transactionCode>S</transactionCode>
                <equitySwapInvolved>0</equitySwapInvolved>
            </transactionCoding>
            <transactionTimeliness>
                <value></value>
            </transactionTimeliness>
            <transactionAmounts>
                <transactionShares>
                    <value>2400444</value>
                </transactionShares>
                <transactionPricePerShare>
                    <value>13.50</value>
                </transactionPricePerShare>
                <transactionAcquiredDisposedCode>
                    <value>D</value>
                </transactionAcquiredDisposedCode>
            </transactionAmounts>
            <postTransactionAmounts>
                <sharesOwnedFollowingTransaction>
                    <value>22805668</value>
                </sharesOwnedFollowingTransaction>
            </postTransactionAmounts>
            <ownershipNature>
                <directOrIndirectOwnership>
                    <value>I</value>
                    <footnoteId id="F1"/>
                    <footnoteId id="F2"/>
                </directOrIndirectOwnership>
                <natureOfOwnership>
                    <value>By limited partnership (2)</value>
                </natureOfOwnership>
            </ownershipNature>
        </nonDerivativeTransaction>
    </nonDerivativeTable>

    <footnotes>
        <footnote id="F1">Shares sold include (i) 1,755,277 shares of common stock owned by Spectrum Equity Investors V, L.P.; (ii) 8,289 shares of common stock owned by Spectrum V Investment Managers' Fund, L.P.; (iii) 626,352 shares of common stock owned by Spectrum Equity Investors III, L.P.; and (iv) 10,526 shares of common stock owned by Spectrum III Investment Managers' Fund, L.P. that the respective Spectrum entities sold as part of Ancestry.com Inc.'s initial public offering, which was completed on November 10, 2009.</footnote>
        <footnote id="F2">Shares of common stock held after sale consists of an aggregate of 22,805,668 shares, including (i) 16,676,204 shares held by Spectrum Equity Investors V, L.P.; (ii) 78,748 shares held by Spectrum V Investment Managers' Fund, L.P.; (iii) 5,950,719 shares held by Spectrum Equity Investors III, L.P.; and (iv) 99,997 shares held by Spectrum III Investment Managers' Fund, L.P.  Mr. Parker is a managing director of the general partner of the general partner of Spectrum Equity Investors V, L.P. and a managing director of the general partner of Spectrum V Investment Managers' Fund, L.P.  Mr. Parker is also a limited partner of the general partner of Spectrum Equity Investors V, L.P., a limited partner of Spectrum V Investment Managers' Fund, L.P., a limited partner of the general partner of Spectrum Equity Investors III, L.P. and a limited partner of Spectrum III Investment Managers' Fund, L.P.  Mr. Parker disclaims beneficial ownership of these shares.</footnote>
    </footnotes>

    <remarks></remarks>

    <ownerSignature>
        <signatureName>Christopher Brookhart, Attorney-in-Fact</signatureName>
        <signatureDate>2009-11-10</signatureDate>
    </ownerSignature>
</ownershipDocument>
</XML>
</TEXT>
</DOCUMENT>
</SEC-DOCUMENT>
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